Contribution to the knowledge of the Carabus (Archiplectes) satyrus Kurnakov, 1962, species complex in Abkhazia (Coleoptera, Carabidae, Carabini)

Abstract

This study is based on a comparative analysis of extensive material of Carabus (Archiplectes) satyrus Kurnakov, 1962, its various forms and related taxa recently collected by the authors and some other collectors in Abkhazia. The status or specific affiliations of several subspecies are changed and a subspecies is described. Carabus (A.) besleticus Kurnakov, 1972, stat. n. is treated as a separate species housing six hitherto established subspecies in addition to the nominal type: C. (A.) besleticus mtsaranus Kurnakov, 1972, C. (A.) besleticus duripshensis Kurnakov, 1972, C. (A.) besleticus napraensis Belousov & Zamotajlov, 1993, C. (A.) besleticus dsychvensis Kurnakov, 1972, C. (A.) besleticus adzinbai Retezár, 2013, and C. (A.) besleticus resheviensis subsp. n. Carabus (A.) satyrus is treated as monotypical while the specific status of C. (A.) pseudopshuensis Zamotajlov, 1991, earlier proposed by Fominykh and Zamotajlov (2012), is confirmed based on the morphological and morphometric data.