Genome-Wide Association Study Identifies <i>CDKN1A</i> as a Novel Locus Associated with Muscle Fiber Composition
Ekaterina A. Semenova,
Hirofumi Zempo,
Eri Miyamoto-Mikami,
Hiroshi Kumagai,
Andrey K. Larin,
Rinat I. Sultanov,
Konstantin A. Babalyan,
Andrey V. Zhelankin,
Takuro Tobina,
Keisuke Shiose,
Ryo Kakigi,
Takamasa Tsuzuki,
Noriko Ichinoseki-Sekine,
Hiroyuki Kobayashi,
Hisashi Naito,
Jatin Burniston,
Edward V. Generozov,
Noriyuki Fuku,
Ildus I. Ahmetov
Affiliations
Ekaterina A. Semenova
Department of Molecular Biology and Genetics, Federal Research and Clinical Center of Physical-Chemical Medicine of Federal Medical Biological Agency, 119435 Moscow, Russia
Hirofumi Zempo
Faculty of Health and Nutrition, Tokyo Seiei College, Tokyo 124-0025, Japan
Eri Miyamoto-Mikami
Graduate School of Health and Sports Science, Juntendo University, Chiba 270-1695, Japan
Hiroshi Kumagai
Graduate School of Health and Sports Science, Juntendo University, Chiba 270-1695, Japan
Andrey K. Larin
Department of Molecular Biology and Genetics, Federal Research and Clinical Center of Physical-Chemical Medicine of Federal Medical Biological Agency, 119435 Moscow, Russia
Rinat I. Sultanov
Department of Molecular Biology and Genetics, Federal Research and Clinical Center of Physical-Chemical Medicine of Federal Medical Biological Agency, 119435 Moscow, Russia
Konstantin A. Babalyan
Department of Molecular Biology and Genetics, Federal Research and Clinical Center of Physical-Chemical Medicine of Federal Medical Biological Agency, 119435 Moscow, Russia
Andrey V. Zhelankin
Department of Molecular Biology and Genetics, Federal Research and Clinical Center of Physical-Chemical Medicine of Federal Medical Biological Agency, 119435 Moscow, Russia
Takuro Tobina
Faculty of Nursing and Nutrition, University of Nagasaki, Nagasaki 851-2195, Japan
Keisuke Shiose
Faculty of Education, University of Miyazaki, Miyazaki 889-2192, Japan
Ryo Kakigi
Faculty of Management & Information Science, Josai International University, Chiba 283-8555, Japan
Takamasa Tsuzuki
Faculty of Pharmacy, Meijo University, Nagoya 468-8503, Japan
Noriko Ichinoseki-Sekine
Graduate School of Health and Sports Science, Juntendo University, Chiba 270-1695, Japan
Hiroyuki Kobayashi
Department of General Medicine, Mito Medical Center, Tsukuba University Hospital, Ibaraki 310-0015, Japan
Hisashi Naito
Graduate School of Health and Sports Science, Juntendo University, Chiba 270-1695, Japan
Jatin Burniston
Research Institute for Sport and Exercise Sciences, Liverpool John Moores University, Liverpool L3 5AF, UK
Edward V. Generozov
Department of Molecular Biology and Genetics, Federal Research and Clinical Center of Physical-Chemical Medicine of Federal Medical Biological Agency, 119435 Moscow, Russia
Noriyuki Fuku
Graduate School of Health and Sports Science, Juntendo University, Chiba 270-1695, Japan
Ildus I. Ahmetov
Department of Molecular Biology and Genetics, Federal Research and Clinical Center of Physical-Chemical Medicine of Federal Medical Biological Agency, 119435 Moscow, Russia
Muscle fiber composition is associated with physical performance, with endurance athletes having a high proportion of slow-twitch muscle fibers compared to power athletes. Approximately 45% of muscle fiber composition is heritable, however, single nucleotide polymorphisms (SNP) underlying inter-individual differences in muscle fiber types remain largely unknown. Based on three whole genome SNP datasets, we have shown that the rs236448 A allele located near the cyclin-dependent kinase inhibitor 1A (CDKN1A) gene was associated with an increased proportion of slow-twitch muscle fibers in Russian (n = 151; p = 0.039), Finnish (n = 287; p = 0.03), and Japanese (n = 207; p = 0.008) cohorts (meta-analysis: p = 7.9 × 10−5. Furthermore, the frequency of the rs236448 A allele was significantly higher in Russian (p = 0.045) and Japanese (p = 0.038) elite endurance athletes compared to ethnically matched power athletes. On the contrary, the C allele was associated with a greater proportion of fast-twitch muscle fibers and a predisposition to power sports. CDKN1A participates in cell cycle regulation and is suppressed by the miR-208b, which has a prominent role in the activation of the slow myofiber gene program. Bioinformatic analysis revealed that the rs236448 C allele was associated with increased CDKN1A expression in whole blood (p = 8.5 × 10−15) and with greater appendicular lean mass (p = 1.2 × 10−5), whereas the A allele was associated with longer durations of exercise (p = 0.044) reported amongst the UK Biobank cohort. Furthermore, the expression of CDKN1A increased in response to strength (p p = 0.00035) training. Accordingly, we found that CDKN1A expression is significantly (p = 0.002) higher in the m. vastus lateralis of strength athletes compared to endurance athletes and is positively correlated with the percentage of fast-twitch muscle fibers (p = 0.018). In conclusion, our data suggest that the CDKN1A rs236448 SNP may be implicated in the determination of muscle fiber composition and may affect athletic performance.