Mature radio galaxies such as M87 belong to a specific subclass of active galaxies (AGN) whose evolution in time endows them with five distinguishing characteristics, including (1) low excitation emission, (2) low star formation rates, (3) high bulge stellar-velocity dispersion, (4) bright stellar nuclei, and (5) weak or nonexistent merger signatures. We show how to understand these seemingly disparate characteristics as originating from the time evolution of powerful radio quasars and describe a new model prediction that tilted accretion disks in AGN are expected to occur in bright quasars but not in other subclasses of AGN. The picture we present should be understood as the most compelling evidence for counter-rotation as a key element in feedback from accreting black holes.