Anatomical notes and discussion of the first described aetosaur Stagonolepis robertsoni (Archosauria: Suchia) from the Upper Triassic of Europe, and the use of plesiomorphies in aetosaur biochronology

Abstract

Stagonolepis robertsoni, from the Late Triassic of Scotland, was the first named aetosaurian. Known mostly from a series of natural molds from two localities, the osteology of this taxon has been difficult to interpret. Detailed work on this material in the late 1950s resulted in a monograph that set the standard for the understanding of aetosaurians, making Stagonolepis robertsoni the best known aetosaurian; however, little has been done with this material since. Reanalysis of this material shows that despite its limitations the early 1960s reconstruction work depicts the preserved characteristics faithfully, especially in the skull. The first cervical rib is extremely anteroposteriorly elongate as in Alligator, a character not previously recognized in aetosaurians. Diapophyseal and zygapophyseal vertebral laminae are present in the cervical and trunk vertebrae. The ilium is autapomorphic with distinct pre- and post-processes of the iliac blade. The osteoderms differ from North and South American material that has been ascribed to the genus. Those assignments are based on plesiomorphies within Aetosauria, such as a radial ornamentation and a posteriorly located and medially offset dorsal eminence. Biostratigraphic correlations using taxonomic conclusions based on plesiomorphic characters should not be used. The holotype specimen of S. robertsoni is currently diagnostic, in part because ventral osteoderms are not known for many aetosaurian taxa and the surface ornamentation of randomly distributed, closely packed oblong pits found in S. robertsoni is unique within Aetosauria.

Keywords

• Triassic
• Aetosauria
• Biostratigraphy
• Apomorphy
• Osteoderms
• Pseudosuchia