Description of the first-instar larva of Geocharidius (Coleoptera: Carabidae: Trechitae) with a discussion of the phylogeny of the subtribe Anillina

Abstract

This paper describes the first-instar larva of Geocharidius Jeannel, a species from Mexico, which is the second record of an Anillina (Coleoptera: Carabidae: Trechitae) larva; previously described was a species of the European genus Typhlocharis. Larvae of these two genera share ten synapomorphic characters, which support the monophyletic origin of Anillina. Sister-group relationships of Anillina with Tachyina + Xystosomina are proposed on the basis of three shared larval synapomorphies: seta LA5 of ligula absent; coronal suture in first-instar larvae very short or absent; second- third-instar larvae have none or one secondary seta on lateral sides of stipes and labium and none on mandibles.

Keywords

• geocharidius
• anillina
• trechitae
• carabidae
• coleoptera
• larva
• description
• phylogeny