Whole-plant corn silage (WPCS) is a major source of forage for dairy cattle in Brazil. Improved kernel processing may be especially advantageous when feeding corn hybrids with vitreous endosperms. Two experiments were conducted to evaluate the effects of the theoretical length of cut (TLOC) and ensiling time on WPCS particle size and kernel processing with two types of forage harvesters. In the first trial, the plant was harvested by a pull-type forage harvester (PTFH) at TLOCs of 3, 6 and 9 mm. In the second trial, the harvesting was performed by a self-propelled forage harvester (SPFH) at TLOCs of 6, 12 and 18 mm, with a roll gap of 3 mm. The WPCS were stored for 0, 35 and 140 days. In the PTFH trial, the TLOCs of 3 and 6 mm did not affect the WPCS particle size distribution and mean particle length (MPL). However, the TLOC of 9 mm increased the MPL. The increase in the TLOC with the SPFH led to a higher MPL and percentage of long particles. The ensiling time increased the MPL and long particles only for the WPCS harvested by the SPFH. The strategy of reducing the TLOC with the SPFH increased the percentage of kernels smaller than 4.75 mm. Furthermore, the TLOC of 6 mm led to the best kernel processing with the SPFH. The ensiling time reduced the particle size of the kernel fractions for both forage harvesters. The corn silage processing score was improved with 140 days of ensiling with the SPFH. These findings suggest that increasing the ensiling time and a low TLOC in SPFHs (6 mm) may be good strategies to increase kernel damage and starch digestibility in WPCS.