New record and range extension of the Horned Toad, Rhinella ceratophrys (Boulenger, 1882) (Anura: Bufonidae), in Venezuela, and confirmation of its presence in Brazil

Abstract

The Horned Toad, Rhinella ceratophrys, is widely distributed in the northwestern part of the Amazon region. It is known from Venezuela by only a single juvenile from the base of Cerro Marahuaca, Amazonas state. Herein we report four additional specimens from Venezuela: three juveniles also from Cerro Marahuaca, and a subadult female from the southern slopes of Cerro La Neblina, at the southern border of Amazonas state. The specimen from Cerro La Neblina extends the species distribution to the extreme south of Venezuelan Amazonia, 307 km south of Marahuaca, and it represents the southeasternmost record of the species. We also confirm the presence of R. ceratophrys in Brazil based on voucher specimens, and present an updated distribution map.