Crying With Laughter: Adapting the Tickling Protocol to Address Individual Differences Among Rats in Their Response to Playful Handling

• Vincent Bombail,
• Sarah M. Brown,
• Tayla J. Hammond,
• Tayla J. Hammond,
• Simone L. Meddle,
• Birte L. Nielsen,
• Birte L. Nielsen,
• Emma K. L. Tivey,
• Alistair B. Lawrence,
• Alistair B. Lawrence

Affiliations

Vincent Bombail

Physiologie de la Nutrition et du Comportement Alimentaire (PNCA), INRAE, Université Paris-Saclay, Paris, France

Sarah M. Brown

The Roslin Institute, The Royal (Dick) School of Veterinary Studies, The University of Edinburgh, Edinburgh, United Kingdom

Tayla J. Hammond

The Roslin Institute, The Royal (Dick) School of Veterinary Studies, The University of Edinburgh, Edinburgh, United Kingdom

Tayla J. Hammond

Animal and Veterinary Sciences, Scotland's Rural College (SRUC), Edinburgh, United Kingdom

Simone L. Meddle

The Roslin Institute, The Royal (Dick) School of Veterinary Studies, The University of Edinburgh, Edinburgh, United Kingdom

Birte L. Nielsen

Modélisation Systémique Appliquée aux Ruminants (MoSAR), INRAE, Université Paris-Saclay, Paris, France

Birte L. Nielsen

Universities Federation for Animal Welfare (UFAW), Wheathampstead, United Kingdom

Emma K. L. Tivey

The Roslin Institute, The Royal (Dick) School of Veterinary Studies, The University of Edinburgh, Edinburgh, United Kingdom

Alistair B. Lawrence

The Roslin Institute, The Royal (Dick) School of Veterinary Studies, The University of Edinburgh, Edinburgh, United Kingdom

Alistair B. Lawrence

Animal and Veterinary Sciences, Scotland's Rural College (SRUC), Edinburgh, United Kingdom